Charline Smadi


I am interested in Probability Theory applied to Ecology and Evolution.
Research interests: branching processes, random environment, population dynamics, population genetics, mate choice.


Publications in Mathematics

  1. C. Smadi, V.A. Vatutin, Critical branching processes in random environment with immigration: survival of a single family, Extremes, 2021 (arXiv:1911.00316).
  2. C. Coron, M. Costa, F. Laroche, H. Leman, C. Smadi, Emergence of homogamy in a two-loci stochastic population model, ALEA, Lat. Am. J. Probab. Math. Stat., 18:469–508, 2021.
  3. C. Dong, C. Smadi, V.A. Vatutin, Critical branching processes in random environment and Cauchy domain of attraction, ALEA, Lat. Am. J. Probab. Math. Stat., 17:877–900, 2020.
  4. A. Gonzalez Casanova, C. Smadi, On Λ-Fleming–Viot processes with general frequency-dependent selection, Journal of Applied Probability, 57(4):1162-1197 (arXiv:1903.06406), 2020.
  5. A. Bovier, L. Coquille, C. Smadi, Crossing a fitness valley as a metastable transition in a stochastic population model, Annals of Applied Probability, 2019 (arXiv:1801.06473).
  6. C. Coron, M. Costa, H. Leman, C. Smadi, A stochastic model for speciation by mating preferences, Journal of Mathematical Biology, 76:1421–1463, 2018.
  7. C. Smadi, The effect of recurrent mutations on genetic diversity in a large population of varying size, Acta Applicandae Mathematicae, 149(1): 11-51, 2017 (arXiv:1511.03538).
  8. S. Billiard, C. Smadi, The interplay of two mutations in a population of varying size: a stochastic eco-evolutionary model for clonal interference, Stochastic Processes and their Applications, 127(3): 701-748, 2017 (arXiv:1505.04550).
  9. R. Brink-Spalink, C. Smadi, Genealogies of two linked neutral loci after a selective sweep in a large population of varying size, Advances in Applied Probability, 49(1): 279-326, 2017 (arXiv:1502.03837).
  10. S. Palau, J.C. Pardo, C. Smadi, Asymptotic behaviour of exponential functionals of Lévy processes with applications to random processes in random environment, ALEA, Lat. Am. J. Probab. Math. Stat., 13, 1235–1258, 2016.
  11. C. Smadi, V.A. Vatutin, Reduced two-type decomposable critical branching processes with possibly infinite variance, Markov Processes and Related Fields, 21(2): 311-358, 2016 (arXiv:1508.06653).
  12. C. Smadi, An Eco-Evolutionary approach of Adaptation and Recombination in a large population of varying size, Stochastic Processes and their Applications, 125(5): 2054 – 2095, 2015, (arXiv:1402.4104).
  13. V. Bansaye, J.C. Pardo, C. Smadi, On the extinction of Continuous State Branching Processes with catastrophes, Electronic Journal of Probability, 18 no. 106, 1-31, 2013.

Publications in Ecology and Evolution

  1. L. Maisonneuve, T. Beneteau, M. Joron, C. Smadi, V. Llaurens, When do opposites attract? A model uncovering the evolution of disassortative mating, to appear in The American Naturalist (biorxiv.2020.05.19.104190).
  2. S. Billiard, C. Smadi, Stochastic dynamics of three competing clones: Conditions and times for invasion, coexistence and fixation , The American Naturalist 195 (3), 463-484, 2020 (arXiv:1607.04656).
  3. F. Lavallée, C. Smadi, I. Alvarez, B. Reineking, F-M. Martin, F. Dommanget, S. Martin, A stochastic individual based model for the growth of a stand of Japanese knotweed including mowing as a management technique, Ecological Modelling, 413, 2019.
  4. T. Cordonnier, C. Smadi, G. Kunstler, B. Courbaud, Asymmetric competition, ontogenetic growth and size inequality drive the difference in productivity between two-strata and one-stratum forest stands, Theoretical Population Biology, 130: 83-93, 2019.
  5. M. Voinson, A. Alvergne, S. Billiard, C. Smadi, Stochastic dynamics of an epidemic with recurrent spillovers from an endemic reservoir, Journal of Theoretical Biology, 457:37-50, 2018.
  6. C. Smadi, H. Leman, V. Llaurens, Looking for the right mate in diploid species: how does genetic dominance affect spatial differentiation in a sexual trait?, Journal of Theoretical Biology, 447:154-170, 2018.


  1. L. Maisonneuve, C. Smadi, V. Llaurens, The limits of evolutionary convergence in sympatry: reproductive interference and developmental constraints leading to local diversity in aposematic signals (biorxiv.2021.01.22.427743)
  2. A. Marguet, C. Smadi, Parasite infection in a cell population with deaths (arXiv:2010.16070).
  3. A. Marguet, C. Smadi, Long time behaviour of continuous-state nonlinear branching processes with catastrophes (arXiv:1908.11592).
  4. C. Coquille, A. Kraut, C. Smadi, Stochastic individual-based models with power law mutation rate on a general finite trait space (arXiv:2003.03452).
  5. V. Bansaye, J.C. Pardo, C. Smadi, Extinction rate of continuous state branching processes in critical Lévy environments, (arXiv:1903.06058).


Isabelle Alvarez, Alexandra Alvergne, Vincent Bansaye, Sylvain Billiard, Anton Bovier, Loren Coquille, Thomas Cordonnier, Camille Coron, Manon Costa, Benoît Courbaud, Fanny Dommanget, Adrian Gonzalez Casanova, Mathieu Joron, Georges Kunstler, Fabien Laroche, Hélène Leman, Violaine Llaurens, Aline Marguet, François-Marie Martin, Sophie Martin, Sandra Palau, Juan Carlos Pardo, Björn Reineking, Vladimir Vatutin